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Skeletons are generally needed for structural support for land organisms in order for their bodies to overcome gravity and aid in locomotion. What are the underlying structural and physical reasons for the evolution of the endoskeleton in vertebrate fish, however? In water there generally tends to be no need to generate a structural support for the purpose of overcoming gravity alone. The density of biological tissue might be slightly greater than water, but swim bladders or the like tend to work relatively well as a counterbalance to those types of forces. For locomotion, you need to have low drag or resistance to movement as you are swimming through the water, but at the same time an organism also needs to impart a backward force to the water itself in order to propel that organism forward. The generation of 'lift' would not necessesarily be as critical to a water organism as it would be to an air organism, again, because of the lesser density differential between the organism itself and its surrounding medium, when an organism is floating in water rather than when it is flying in air. You generally tend to need a situation where the organism can generate a high water resistance, impart a force, then generate a lower water resistance in order to be impelled forward while simultaneously extending itself back to its initial conditions so that it can impart a new force, all without producing an opposite backward force of the same magnitude, which would, ultimately, make it go nowhere if were not minimised. Now it just seems to me that fish tend to swim faster than jellyfish, although I can't say for certain. The early chordates were supposed to have a sessile form and a younger moving form, that gradually became the fishes. It would seem to me that a notochord or backbone would enable a sessile organism to thrive in rapidly flowing waters withoug being torn apart or rupturing. Nonetheless, the notochord tends to be a structure for the larval rather than the adult state for at least some of the most primitave chordates, although I am not sure of the relative extents. Are there any structural reasons with underlying physics involved, as to why a backbone or notochord might be advantageous for the purpose of locomotion speeds in the early fishes, rather than simply having opposing muscles resist contraction on the opposite side through closed water chambers, or the like? What advantage does a fish have, in comparison to a swimming worm, an octopus, or a starfish?
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